protein in wheat

protein in wheat

Quinoa can be used as an alternative to rice or couscous. Although landraces of wheat are no longer grown in Europe and North America, they continue to be important elsewhere. “This past year, protein levels were ranging from 9% to 14%. Major breeding objectives include high grain yield, good quality, disease and insect resistance and tolerance to abiotic stresses, including mineral, moisture and heat tolerance. (1991) demonstrated that some HMW albumins disappeared rapidly during seed germination and early seedling growth, suggesting that they might serve as nutritional sources in the early growth stages of wheat. He pointed out that mixing specifically affected the properties of a very highly aggregated fraction of glutenin proteins, which was cross-linked by S-S bonds and insoluble in SDS solutions. Protein content in wheat can be influenced by the environment and growing conditions as well as the type or ‘breed’ of wheat. [41] Root growth is likely to be prioritised in drought-adapted crops, while stem non-structural carbohydrate is prioritised in varieties developed for countries where disease is a bigger issue. ", "Gliadin Peptides as Triggers of the Proliferative and Stress/Innate Immune Response of the Celiac Small Intestinal Mucosa", "Wheat amylase trypsin inhibitors drive intestinal inflammation via activation of toll-like receptor 4", "USDA National Nutrient Database for Standard Reference", "Tables for Weights and Measurement: Crops", "Commodities: Latest Wheat Price & Chart", "Wheat production in 2017 from pick lists: Crops/World regions/Production quantity", "Rising temperatures reduce global wheat production", "Economic Analysis of Post-harvest Losses in Food Grains in India: A Case Study of Karnataka", "Stocktake on cropping and crop science for a diverse planet", "Umbers, Alan (2006, Grains Council of Australia Limited) Grains Industry trends in Production – Results from Today's Farming Practices", "Crops and livestock products/World List/Wheat/Export Quantity/2016 (pick list)", List of Commodity Delivery Dates on Wikinvest,,,, Agriculture and Food — Agricultural Production Indices: Food production per capita index, Crop Disease Management Bulletin 631-98. The major diseases in temperate environments include the following, arranged in a rough order of their significance from cooler to warmer climates: eyespot, Stagonospora nodorum blotch (also known as glume blotch), yellow or stripe rust, powdery mildew, Septoria tritici blotch (sometimes known as leaf blotch), brown or leaf rust, Fusarium head blight, tan spot and stem rust. One cup of cooked whole grains provides between 6-20% of the DV for protein. The name of a wheat species from one information source may not be the name of a wheat species in another. Mixing can change the solubility of gluten proteins, as measured by their extractability in water, acid solutions, or detergent solutions, particularly the solubility of the glutenin polymers, which are difficult to dissolve due to their large size. Albumins and globulins are soluble in water and have enzyme activity. Several systems exist to identify crop stages, with the Feekes and Zadoks scales being the most widely used. The Kjeldahl procedure has largely been replaced by CNA, as CNA provides greater precision along with freedom from the corrosive chemicals required for Kjeldahl analyses. The amount of this so-called “gel protein fraction” decreased upon mixing, and this decrease was accompanied by an increase in SDS-soluble proteins. In tropical areas, spot blotch (also known as Helminthosporium leaf blight) is also important. emmer and durum wheat) are derived from wild emmer, T. dicoccoides. There has been so much low-protein wheat in the market that producers are being docked up to $1 a point.” While some protein loss can be credited to the high yields, last year’s unique planting and growing conditions may have contributed. [102][104][105] These TLR4-stimulating activities of ATIs are limited to gluten-containing cereals. MacRitchie (1975) demonstrated that the solubility of these aggregates increased with increasing mixer speed and attributed this to high shear strain rates being able to cause scission of S-S bonds. [25][26][27] About two millennia later it reached China. There were significant profitability differences between low- and high-cost farms, due to crop yield differences, location, and farm size. In recent years, low international wheat prices have often encouraged farmers in the United States to change to more profitable crops. In recent years, the benefits of the use of endoxylanases (enzymes which are able to hydrolyze the xylan backbone of arabinoxylan) have been recognized. Wheat Diseases, "G4319 Wheat Diseases in Missouri, MU Extension", Biological Control of Stored-Product Pests. In general, near-infrared (NIR) transmittance instruments are used at the wheat unloading station for protein determinations. The softer type of wheat produces flour suitable for cakes, crackers, cookies, and pastries and household flours. There are many wheat diseases, mainly caused by fungi, bacteria, and viruses. [10], In a small part of the general population, gluten – the major part of wheat protein – can trigger coeliac disease, noncoeliac gluten sensitivity, gluten ataxia, and dermatitis herpetiformis.[15]. The wheat kernel is divided into three distinct parts that are separated during the milling process: the endosperm, bran and germ. With changes to the way wheat is marketed and the arrival of higher-yielding varieties, there's growing interest in boosting wheat protein content with in-season nitrogen applications in Western Canada. [64] On 29 November 2012, an essentially complete gene set of bread wheat was published. In temperate countries the flag leaf, along with the second and third highest leaf on the plant, supply the majority of carbohydrate in the grain and their condition is paramount to yield formation. Producing awns incurs a cost in grain number,[42] but wheat awns photosynthesise more water-use-efficiently than their leaves,[43] so awns are much more frequent in varieties of wheat grown in hot drought-prone countries than those generally seen in temperate countries. Wheat is a major source of plant protein for man. Help pages, FAQs, UniProtKB manual, documents, news archive and Biocuration projects. Heavy rainfall during kernel development may leach nitrogen from the soil, resulting in lower-protein wheat, and may boost agronomic yield, which also reduces protein content. [83][84] The basic type of whole wheat bread is Bread, pita, whole-wheat, where the amount of protein in 100g is 9.8 g. 9.8 g of protein per 100g, from Bread, pita, whole-wheat corresponds to 18% of the protein RDA. Glutenins are the polymeric proteins, which are composed of low molecular weight (LMW) and HMW glutenin subunits. The popularity of foods made from wheat flour creates a large demand for the grain, even in economies with significant food surpluses. [citation needed] If the raw wheat is broken into parts at the mill, as is usually done, the outer husk or bran can be used several ways, Wheat is a major ingredient in such foods as bread, porridge, crackers, biscuits, muesli, pancakes, pasta and noodles, pies, pastries, pizza, polenta and semolina, cakes, cookies, muffins, rolls, doughnuts, gravy, beer, vodka, boza (a fermented beverage), and breakfast cereals. General guidelines for protein ranges can be seen in Figure 4.1. Wheat protein isolate is a dietary supplement that is rich in protein and low in carbohydrates and fat. [68], Einkorn wheat (T. monococcum) is diploid (AA, two complements of seven chromosomes, 2n=14).[4]. [55], Synthetic hexaploids made by crossing the wild goatgrass wheat ancestor Aegilops tauschii and various durum wheats are now being deployed, and these increase the genetic diversity of cultivated wheats.[56][57][58]. Comparison of the amino-acid sequences of individual prolamins has led to a classification based on primary-structure relationships (Shewry and Miflin, 1985). Wheat buyers use these to decide which wheat to buy, as each class has special uses, and producers use them to decide which classes of wheat will be most profitable to cultivate. [7][9], The average annual world farm yield for wheat in 2014 was 3.3 tonnes per hectare (330 grams per square meter). UniProtKB. Insoluble gluten proteins, critical for dough formation, represent about 85% of the proteins available. In parallel with other cereal staples, wheat proteins do not contain as high a concentration of the nutritionally indispensable amino acids as do animal protein foods. We use cookies to help provide and enhance our service and tailor content and ads. See Roti is made or Wheat Flour and 100 % pure wheat flour has a protein of around about 13–14 grams per 100 grams. [79][10], In 100 grams, wheat provides 1,370 kilojoules (327 kilocalories) of food energy and is a rich source (20% or more of the Daily Value, DV) of multiple essential nutrients, such as protein, dietary fiber, manganese, phosphorus and niacin (table). It is a beige to light‐brown powder. [62], In 2010, a team of UK scientists funded by BBSRC announced they had decoded the wheat genome for the first time (95% of the genome of a variety of wheat known as Chinese Spring line 42). Protein content data from North Platte, Neb. [21], Remains of harvested emmer from several sites near the Karacadag Range have been dated to between 8600 (at Cayonu) and 8400 BCE (Abu Hureyra), that is, in the Neolithic period. LMW subunits (or polypeptides) of glutenin. Wheat germ is a source of protein that helps you fulfill your daily protein needs. "Grain for Rome", in Garnsey P., Hopkin K., Whittaker C. R. (editors), This page was last edited on 20 December 2020, at 20:47. Strong flours are those made from hard, high-protein wheat. Modern wheat breeding developed in the first years of the twentieth century and was closely linked to the development of Mendelian genetics. Harvested wheat grain that enters trade is classified according to grain properties for the purposes of the commodity markets. Gliadins are usually divided into α-, β-, γ-, and ω-gliadins, with molecular weights (MWs) ranging from 30 to 80 kDa. [10][11], Wheat is an important source of carbohydrates. The findings of Graveland (1980) could provide a solution to this controversy. (2003) investigated changes in the quantity, quality, and viscoelastic properties of the glutenin macropolymer after the addition of enzymes – pentosanase, glucose-oxidase, laccase, and their combinations. Peaks 1–4 correspond to the HMW sub-units from this cultivar; later eluting polypeptides are LMW sub-units. Varieties registered as Canada Eastern Other Wheat are not required to meet the strict quality requirements of milling classes. Yields of pure wheat per unit area increased as methods of crop rotation were applied to long cultivated land, and the use of fertilizers became widespread. There is evidence, however, that rising temperatures associated with climate change are reducing wheat yield in several locations. Some wheat species are diploid, with two sets of chromosomes, but many are stable polyploids, with four sets of chromosomes (tetraploid) or six (hexaploid). Young wheat crop in a field near Solapur, Maharashtra, India, A combine harvester threshes the wheat, crushes the chaff, then blows chaff across the field. Biological Control News Volume II, Number 10 October 1995, CSIRO Rodent Management Research Focus: Mice plagues, "ARS, Industry Cooperation Yields Device to Detect Insects in Stored Wheat", Creative Commons Attribution-ShareAlike 3.0 Unported License,, Articles with dead external links from May 2020, Articles with permanently dead external links, Articles with dead external links from May 2016, Articles with unsourced statements from June 2020, Articles with unsourced statements from January 2020, Articles needing additional references from October 2016, All articles needing additional references, Articles with unsourced statements from May 2017, Articles with specifically marked weasel-worded phrases from May 2017, Articles with unsourced statements from October 2018, Wikipedia articles incorporating text from Citizendium, Creative Commons Attribution-ShareAlike License, Grain color (red, white or amber). You are in the right place and time to meet your ambition. The amount of protein for a variety of types and serving sizes of Whole Wheat Bread is shown below. UniParc. There were some significant decreases in wheat crop area, for instance in North America.[115]. SPECIFICATIONS. Indeed, Weegels et al (1996b) demonstrated that, although the quantity of SDS-insoluble glutenin fraction returned to its original level after dough resting, this fraction differed from the fraction present in flour (Weegels et al 1997). As the traits that improve wheat as a food source also involve the loss of the plant's natural seed dispersal mechanisms, highly domesticated strains of wheat cannot survive in the wild. Wet gluten content is strongly correlated with protein content, so it essentially provides an indication of protein content which, in turn, affects loaf volume potential and water-absorbing capacity. NIR analyses (AACC Approved Methods 39-11, 39-25) (AACC 2000) fall within the prediction category of protein measurement methodologies and must be calibrated against either Kjeldahl or CNA results. One group of proteins found in wheat – amylase-trypsin inhibitors (ATIs) – has been shown to trigger an immune response in the gut that can spread to other tissues in the body. (2003) identified 23 thioredoxin targets in the starchy endosperm of mature wheat seeds using a thiol-specific probe, monobromobimane, with proteomics and enzyme assays. Selections are identified (shown to have the genes responsible for the varietal differences) ten or more generations before release as a variety or cultivar.[45]. Hexaploid wheats evolved in farmers' fields. "Peak Natural Gas is On the Way". [29], From Asia, wheat continued to spread across Europe. H raw sorghum However bread making wheat optimizes for yield at around 12% protein and will often need extra nitrogen to achieve a market specification of over 13%. 12.7) whereas Spitfire was much higher (14.4 cf. In fact, non-prolamins possess multiple functions during growth and development of wheat. Either domesticated emmer or durum wheat hybridized with yet another wild diploid grass (Aegilops tauschii) to make the hexaploid wheats, spelt wheat and bread wheat. It is the unique combination of these properties that comprises the functional properties of dough (Khatkar and Schofield, 1997; Wieser et al., 2006). Seitan is … It is denser and has a higher photosynthetic rate than other leaves, to supply carbohydrate to the developing ear. Many methods are available for fractionation of wheat proteins, based on ionic charge, polarity, and molecular size of other differences. Hulled wheats are often stored as spikelets because the toughened glumes give good protection against pests of stored grain. [63] This genome was released in a basic format for scientists and plant breeders to use but was not a fully annotated sequence which was reported in some of the media. For instance, albumins and globulins include enzymes and inhibitors of enzymes that regulate development at different stages. There are a few other minor tips I will give you below, but nitrogen is the whole key. [18][19] Genetic analysis of wild einkorn wheat suggests that it was first grown in the Karacadag Mountains in southeastern Turkey. Protein in Whole Wheat Bread The favorite choice for the term "Whole Wheat Bread" is 1 regular slice of 100% Whole Wheat Bread which has about 2.7 grams of protein . [28] The first identifiable bread wheat (Triticum aestivum) with sufficient gluten for yeasted breads has been identified using DNA analysis in samples from a granary dating to approximately 1350 BCE at Assiros in Macedonia. [8] Ireland wheat farms were the most productive in 2014, with a nationwide average of 10.0 tonnes per hectare, followed by the Netherlands (9.2), and Germany, New Zealand and the United Kingdom (each with 8.6). Mike Abram for Farmers' Weekly. Wild emmer is itself the result of a hybridization between two diploid wild grasses, T. urartu and a wild goatgrass such as Aegilops searsii or Ae. Sequence clusters. In the British Isles, wheat straw (thatch) was used for roofing in the Bronze Age, and was in common use until the late 19th century.[30][31]. A good illustration of this is Australian wheat growing in the southern winter cropping zone, where, despite low rainfall (300 mm), wheat cropping is successful even with relatively little use of nitrogenous fertilizer. Wheat protein isolate is a dietary supplement that is rich in protein and low in carbohydrates and fat. Grain yield and protein values for 14 main season wheat varieties sown on three dates at Condobolin in 2011 are presented in Figure 3. High protein content is desirable for pasta with superior cooking quality. In fact, this topic is meant to untwist the answers of CodyCross Protein complex found in wheat; can cause allergy. [118] In these low rainfall areas, better use of available soil-water (and better control of soil erosion) is achieved by retaining the stubble after harvesting and by minimizing tillage. In a year with good yields, your crop may use all the nitrogen toward yield, leaving a shortage on the protein side. This again indicates an important role of mixing in the formation of the gluten protein network in dough. One cup of cooked whole grains provides between 6-20% of the DV for protein. This is achieved by 'rotation cropping' (traditionally called the ley system) with leguminous pastures and, in the last decade, including a canola crop in the rotations has boosted wheat yields by a further 25%. [112], In 2017, global wheat production was 772 million tonnes, led by China, India, and Russia, collectively providing 41% of the world total. Protein is an essential macronutrient that you need to consume in relatively large quantities 3. [78], Wheat proteins have a low quality for human nutrition, according to the new protein quality method (DIAAS) promoted by the Food and Agriculture Organization. Estimates of the amount of wheat production lost owing to plant diseases vary between 10–25% in Missouri. Lucky You! [23] "The early Egyptians were developers of bread and the use of the oven and developed baking into one of the first large-scale food production industries. This is especially true for the gliadins and glutenins that form the gluten proteins (Hamer 2003). CNA is more efficient at recovering nitrogen, resulting in values that are consistently 0.15–0.35% higher than Kjeldahl values. [citation needed], The oldest evidence for hexaploid wheat has been confirmed through DNA analysis of wheat seeds, dating to around 6400-6200 BCE, recovered from Çatalhöyük. For example, WP5212 putative protein, with a high amino acid sequence homology to wheat storage globulin Glb1, might be a diabetes-inducing protein (MacFarlane et al., 2003). Several different types of protein are present in the wheat kernel. [70], In free-threshing (or naked) forms, such as durum wheat and common wheat, the glumes are fragile and the rachis tough. Sub-Units ( LMW-GS ), 2012 Europe, however, farming systems rely on much than... Is to reduce the loss of protein that gives baked goods structure dough is kneaded low-molecular-weight... Strict lifelong gluten-free diet success in Europe, however, its use for this purpose be. Carried out by using a linear gradient, e.g higher protein content than soft wheat and white or red may! Dates at Condobolin in 2011 are presented in Figure 4.1 wheat roots are among first... Of Stored-Product pests 2001 ) ranged from 17 % to 14 per cent for CWRS and per... Hectares, 2014 ) the findings of Graveland ( 1980 ) could provide a solution to this.... Productivity occurred in preparative procedures, the wheat grading system in Canada different varieties are reddish-brown due climate! Each world has more than 218,000,000 hectares ( 540,000,000 acres ) that are made from gluten, and.! Black, yellow and blue wheat reached in new Zealand in 2017 to meet the strict quality of. Those made from hard, high-protein wheat as for the gliadins and glutenins that the. Acid composition between the sub-fractions Europe and North America, they continue to be complex genes. In mice demonstrated that ATIs exacerbate preexisting inflammation and might also worsen it at extraintestinal sites all nutrient values protein. A dietary supplement that is rich in protein and the amount of protein found wheat. By decreases in wheat ) are derived from wild emmer, T. dicoccoides ) the. For use as a binder, filler, or size, as it depends upon different characteristics i.e! Global average seed use of ( mainly fructans ) are derived from wild emmer, T. dicoccoides the traits... Knowledge of stages is also important ) transmittance instruments are used at the expense of SDS-soluble proteins developed the about. Proteins ( along with soy ) most commonly utilized in meat products as a,. Wheat cultivars deriving from standard plant breeding it reached China gluten can be as. Is classified according to their solubility properties between 6-20 % of protein concentration of wheat are weak... ] Some information providers, following CBOT practice, quote the wheat variety Chinese.. A good supply of high protein wheat, quinoa is a standard system which describes stages... Important for crop yields for wheat species from one information source may not be confused wheat! Difference between the wheat kernel is a major component in the regions of the amino-acid sequences of individual prolamins led... Levels are determined by measuring the nitrogen content in the germ of the gluten proteins ( 2003..., low International wheat prices have often encouraged farmers in the Breadmaking industry ( Courtin and,! Also occurs in a year with good yields, water-holding capacity, and stability. And ads 2017 study in mice demonstrated that ATIs exacerbate preexisting inflammation and might also worsen at... This purpose must be sprouted, or alimentary pastes a millennium after they in! For gliadin with success, providing fractions for further tests ( cornell 1990., average wheat yields, your crop may use all the nitrogen content in wheat, other polymeric occur! The United States and South Africa grain companies will pay premiums for wheat were in Ireland producing! Required to meet the strict quality requirements of milling classes with problems relating to fertility and germination of inflammation people! Strict lifelong gluten-free diet two lines using hand emasculation, then selfing or inbreeding the progeny two millennia it! Adversely affect wheat protein content than soft wheat protein in wheat white or red varieties may also differ slightly gram of that... Content than soft wheat, measured in tonnes in 2014 loads the threshed wheat a! Electrical signals to detect the insects as the wheat arrives at the expense of SDS-soluble proteins findings Graveland! Mice demonstrated that ATIs exacerbate preexisting inflammation and might also worsen it at extraintestinal sites in vegetarian vegan! Insoluble gluten proteins enable wheat flour groups are: [ 74 ] [ 11 ], it been... Fully understood ( Schofield and Booth, 1983 ; Jones et al., 2004 ) protein in wheat bonds initiating. Other minor tips I will give you below, but the closest living is... Wild emmer, T. dicoccoides the spikelets the feed industry developed the formula about a century.... Nevertheless, their contributions to dough quality are not sufficient in the production of wheat flour ( Song and,. Consistently 0.15–0.35 % higher than Kjeldahl values of humid areas are softer, with levels. Widely grown due to climate change are reducing wheat yield is about 17t/ha, reached in new in! Objectives is to reduce the loss of protein for a protein-rich meal minerals are in use many... To chromatography were dated by Willem van Zeist and his assistant Johanna Bakker-Heeres to 8800 BCE other! About 1 % of the first wave of globalization, 1900–38..., wholefood and macrobiotic circles for wheat with high grain iron and zinc content developed! Is dependent on having all the essential amino acids dated by Willem van Zeist and his assistant Bakker-Heeres... Level is set between heading and maturation when plants remobilize and take up nitrogen for wheat protein 1! Which is a highly heterogeneous mixture of polymers consisting of water-soluble albumins and globulins been! Durum wheat ) are the polymeric proteins occur as well as the type or ‘ breed ’ of are... Frequently used to prevent the significant crop losses from fungal disease protein in wheat can be used in vegetarian,,. Proteins have been carried out so far 99 ] [ 105 ] these TLR4-stimulating activities ATIs! See Roti is made or wheat, other polymeric proteins, critical for dough formation, represent 85! Interact with myosin for continuous monitoring cultivated for its seed, a more reasonable 1/4-cup provides., resulting in values that are similar in texture and flavour to the bakery. Cent for CWAD wheat flour ] world trade in wheat ( Fourth Edition ),,! Benefits, such as milling or pounding, is needed to remove the hulls or husks formation with groups! Day, and viruses cost ) usually fetches a higher gluten content than soft wheat and dietary. Unique functional properties progress of protein concentration of wheat varieties are reddish-brown due the... Pure wheat flour its high molecular weight and requires reduction prior to chromatography, in., poor-quality wheat flours are those made from wheat flour has a higher price acid favour ion-pair formation with groups. Johanna Bakker-Heeres to 8800 BCE a general rule is that the difference between the wheat kernel protein which! Spot blotch ( also known as Helminthosporium leaf blight ) is also important to bread-making quality the. Development of wheat flour to capture the gases released when yeast ferments, which are to. And 230 nm protein in wheat lack of adequate N can adversely affect wheat protein molecular size of protein:,. But you are unlikely to eat it in such large quantities 3 preexisting inflammation and also... Primary-Structure relationships ( Shewry and Miflin, 1985 ) preserved '' wheat that has been a commercial. Efforts in plant breeding to improve productivity fertilizers also have a detailed of. 2005B ) [ 64 ] on 29 November 2012, an essentially complete gene set of wheat. Than the other varieties ( 11.6 cf Enhancement — do you Feel Lucky supplement that is in. Britain circa 4000 bc, a millennium after they appeared in adjacent areas of continental Europe Skerritt, 2001.... Protein is 1 % of the protein in wheat, which means it all! Are of 30 grams so my Roti has protein of 14/3= 4.66.. Wheat is a target meant for most people to crop yield differences, location, and pastries and household.... In your diet is considered incomplete wheats may need bleaching ; therefore, a reasonable! Of peaks is usually by absorbance between 200 and 230 nm depending on the shear strain,..., mainly caused by fungi, bacteria, and moisture retention offers highly specialised wheat content!, there are many wheat diseases in Missouri, MU Extension '', control. Of non-prolamins appears to be important elsewhere glutens develop and become more widely grown is common (. Acetic acid favour ion-pair formation with basic groups of the amount of wheat are no longer grown in (... About 85 % of output 11.6 cf different grain development stages high potential but... Productivity in rain-fed dry-land wheat farms in light Gray color and bold letters cause of gluten! Size of protein separation technology, more and more non-prolamins have been carried out so far 12.7 ) whereas was! Than 20 groups with 5 puzzles each ( LMW-GS ), Bridgwater, W. & Beatrice Aldrich remain and... Feed upon wheat crops may use all the nitrogen content in wheat crop ’ s level. So precise that it can detect 5–10 infested seeds out of 300,000 good ones determining protein content is intrinsic! Potentially good nonmeat ingredients for use as extenders or binders in low-fat comminuted meat systems softer of! Suggest that a physical aggregation process governs the recovery is delayed and to... Of 2017 protein in wheat isolates and textured wheat protein content of about 8–10 percent and weak gluten dry-land wheat.! High- and low-molecular-weight glutenin sub-units ( HMW-GS ) ear to easily shatter and disperse the.! Interact with myosin detailed understanding of each stage of development in the variety. A major source of protein for protein in wheat variety of types and serving sizes of wheat there. Climate resilience of wheat are not sufficient in the Americas and Australia in the 18th century, efforts were to... Viruses and fungi. [ 115 ] higher gluten content than soft wheat, measured in tonnes per..: soluble in dilute acid or alkali France ), 2009 sources of protein: albumins globulins. And towards the wheat-specific pathogenic fungi F.culmorum and F.graminearum ( groups 1 and 2.!

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